{"id":320,"date":"2017-09-04T14:09:44","date_gmt":"2017-09-04T17:09:44","guid":{"rendered":"http:\/\/www.idstudio.art.br\/clientes\/agasscom\/?page_id=320"},"modified":"2017-11-13T00:22:44","modified_gmt":"2017-11-13T02:22:44","slug":"genetica","status":"publish","type":"page","link":"https:\/\/agasscom.org\/site\/pt\/regulamentos\/genetica\/","title":{"rendered":"Gen\u00e9tica"},"content":{"rendered":"<h2>As Leis de Mendel ou Mendelismo na Cria\u00e7\u00e3o de Agapornis<\/h2>\n<p>Os princ\u00edpios de gen\u00e9tica que hoje conhecemos e aplicamos em nossa cria\u00e7\u00e3o de Agapornis foram descobertos por um monge de Ordem de Santo Agostinho, nascido em 1822 na Sil\u00e9sia: Gregor Johann Mendel.<\/p>\n<p>Mendel, com seu trabalho \u201cExperi\u00eancias com hibridiza\u00e7\u00e3o de plantas\u201d formulou os princ\u00edpios b\u00e1sicos de transmiss\u00e3o de heran\u00e7a e que hoje, embora sabidamente outros mecanismos podem e est\u00e3o envolvidos, os princ\u00edpios de sua teoria continuam sendo ensinados em todo o mundo como LEIS DE MENDEL.<\/p>\n<h3>1\u00aa Lei de Mendel<\/h3>\n<p>Os conceitos de Dominante e Recessivo, vistos anteriormente, s\u00e3o essenciais para que os princ\u00edpios de Mendel sejam entendidos nos quais inicialmente trabalharemos com linhagens puras.<\/p>\n<blockquote><p>\u201cLinhagem \u00e9 o conjunto de descendentes de um ancestral comum.\u201d<\/p><\/blockquote>\n<p>Traduzindo, s\u00e3o os filhos , netos, bisnetos, etc originados de um \u00fanico casal ou ave. Os Agapornis, atualmente, dificilmente se encontram em linhagens puras devido ao grande grau de cruzamentos realizados; isto n\u00e3o quer dizer que sejam mesti\u00e7os, mas que os indiv\u00edduos possuem v\u00e1rios genes mutantes, isto \u00e9; s\u00e3o portadores de diversas muta\u00e7\u00f5es que foram sendo introduzidas no decorrer de gera\u00e7\u00f5es seguidas. Isto quer dizer eu quando compramos um Agapornis, muito dificilmente ele vai transmitir uma \u00fanica cor somente, pois deve ser filho, neto, bisneto,etc de outros de cores diferentes que podem estar \u201cescondidas\u201d.<\/p>\n<blockquote><p>\u201cPor linhagem pura entende-se que apenas um par de genes mutantes est\u00e1 presente.\u201d<\/p><\/blockquote>\n<p>Assim trabalharemos somente com linhagens puras: os past\u00e9is somente transmitir\u00e3o genes past\u00e9is a seus descendentes, os amarelos somente amarelos e assim por diante.<\/p>\n<p>Os descendentes dos cruzamentos efetuados recebem a denomina\u00e7\u00e3o F1 (primeira gera\u00e7\u00e3o), F2 (segunda gera\u00e7\u00e3o), etc.<\/p>\n<table>\n<caption>Exemplo 1<\/caption>\n<thead>\n<tr>\n<th style=\"text-align: center;\"><\/th>\n<th style=\"text-align: center;\" colspan=\"2\">VERDE<\/th>\n<th>\u00d7<\/th>\n<th style=\"text-align: center;\" colspan=\"2\">ARLEQUIM RECESSIVO<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\" colspan=\"2\">P<sup>au<\/sup>P<sup>au<\/sup><\/td>\n<td><\/td>\n<td style=\"text-align: center;\" colspan=\"2\">p<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\">50%<\/td>\n<td style=\"text-align: center;\">50%<\/td>\n<td><\/td>\n<td style=\"text-align: center;\">50%<\/td>\n<td style=\"text-align: center;\">50%<\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">G:<\/th>\n<td style=\"text-align: center;\">P<sup>au<\/sup><\/td>\n<td style=\"text-align: center;\">P<sup>au<\/sup><\/td>\n<td><\/td>\n<td style=\"text-align: center;\">p<sup>au<\/sup><\/td>\n<td style=\"text-align: center;\">p<sup>au<\/sup><\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">F1:<\/th>\n<td style=\"text-align: center;\" colspan=\"2\">P<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<td><\/td>\n<td style=\"text-align: center;\" colspan=\"2\">P<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<\/tr>\n<\/tbody>\n<tfoot>\n<tr>\n<td style=\"text-align: center;\" colspan=\"6\">100% Verde (verde portador de arlequim recessivo)<\/td>\n<\/tr>\n<\/tfoot>\n<\/table>\n<p>Acima vimos que cada um dos pais doa um (1) de seus genes (G) originando seus descendentes \u201cP<sup>au<\/sup>p<sup>au<\/sup>\u201d: verdes heterozigotos para arlequim recessivo.<\/p>\n<p>IMPORTANTE: A cor verde \u00e9 um produto esquizocr\u00f4mico de intera\u00e7\u00e3o de v\u00e1rios genes. O gene \u201cP<sup>au<\/sup>\u201d N\u00c3O \u00e9 respons\u00e1vel pela cor verde, mas apenas UM dos respons\u00e1veis, e aqui nota apenas a aus\u00eancia do fator arlequim recessivo.<\/p>\n<table>\n<caption>Exemplo 2<\/caption>\n<thead>\n<tr>\n<th style=\"text-align: center;\"><\/th>\n<th style=\"text-align: center;\" colspan=\"2\">VERDE\/ARLEQUIM RECESSIVO<\/th>\n<th>\u00d7<\/th>\n<th style=\"text-align: center;\" colspan=\"2\">VERDE\/ARLEQUIM RECESSIVO<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\" colspan=\"2\">P<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<td><\/td>\n<td style=\"text-align: center;\" colspan=\"2\">P<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\">50%<\/td>\n<td style=\"text-align: center;\">50%<\/td>\n<td><\/td>\n<td style=\"text-align: center;\">50%<\/td>\n<td style=\"text-align: center;\">50%<\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">G:<\/th>\n<td style=\"text-align: center;\">P<sup>au<\/sup><\/td>\n<td style=\"text-align: center;\">p<sup>au<\/sup><\/td>\n<td><\/td>\n<td style=\"text-align: center;\">P<sup>au<\/sup><\/td>\n<td style=\"text-align: center;\">p<sup>au<\/sup><\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">F2:<\/th>\n<td style=\"text-align: center;\">P<sup>au<\/sup>P<sup>au<\/sup><\/td>\n<td style=\"text-align: center;\">P<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<td><\/td>\n<td style=\"text-align: center;\">P<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<td style=\"text-align: center;\">p<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\">25%<\/td>\n<td style=\"text-align: center;\" colspan=\"3\">50%<\/td>\n<td style=\"text-align: center;\">25%<\/td>\n<\/tr>\n<\/tbody>\n<tfoot>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\">verde homozigoto<br \/>\nn\u00e3o arlequim recessivo<\/td>\n<td style=\"text-align: center;\" colspan=\"3\">verde heterozigoto<\/td>\n<td style=\"text-align: center;\">arlequim recessivo homozigoto<br \/>\npara arlequim recessivo<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\" colspan=\"2\">75% verde (fen\u00f3tipo)<\/td>\n<td><\/td>\n<td style=\"text-align: center;\" colspan=\"2\">25% pastel (fen\u00f3tipo)<\/td>\n<\/tr>\n<\/tfoot>\n<\/table>\n<table>\n<thead>\n<tr>\n<th style=\"text-align: center;\">fen\u00f3tipo<\/th>\n<th style=\"text-align: center;\">combina\u00e7\u00e3o g\u00eanica<\/th>\n<th style=\"text-align: center;\">freq\u00fc\u00eancia<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td>verde<\/td>\n<td>0,5P<sup>au<\/sup> \u00d7 0,5P<sup>au<\/sup><\/td>\n<td>0,25=25%<\/td>\n<\/tr>\n<tr>\n<td>verde \/ arlequim recessivo<\/td>\n<td>2 ( 0,5P<sup>au<\/sup> \u00d7 0,5p<sup>au<\/sup> )<\/td>\n<td>0,25=25%<\/td>\n<\/tr>\n<tr>\n<td>arlequim recessivo<\/td>\n<td>0,5p<sup>au<\/sup> \u00d7 0,5p<sup>au<\/sup><\/td>\n<td>0,25=25%<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>Observamos uma propor\u00e7\u00e3o de 3:1 (verde: arlequim recessivo) que \u00e9 conseq\u00fc\u00eancia da segrega\u00e7\u00e3o dos genes dos heterozigotos. Esta segrega\u00e7\u00e3o ocorre regularmente, em se tratando de linhagens puras, devido \u00e0 separa\u00e7\u00e3o dos cromossomos.<\/p>\n<p>Esta \u00e9 a 1\u00aa LEI DE MENDEL, tamb\u00e9m conhecida como \u201clei da segrega\u00e7\u00e3o ou da pureza dos gametas\u201d e pode ser definida assim:<\/p>\n<blockquote><p>\u201cOs pares de genes se segregam na forma\u00e7\u00e3o dos gametas e cada gene \u00e9 transmitido em uma frequ\u00eancia de 50% a seus descendentes\u201d<\/p><\/blockquote>\n<table>\n<caption>Exemplo 3<\/caption>\n<thead>\n<tr>\n<th style=\"text-align: center;\"><\/th>\n<th style=\"text-align: center;\" colspan=\"2\">VERDE\/ARLEQUIM RECESSIVO<\/th>\n<th>\u00d7<\/th>\n<th style=\"text-align: center;\" colspan=\"2\">ARLEQUIM RECESSIVO<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\" colspan=\"2\">P<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<td><\/td>\n<td style=\"text-align: center;\" colspan=\"2\">p<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">G:<\/th>\n<td style=\"text-align: center;\">P<sup>au<\/sup><\/td>\n<td style=\"text-align: center;\">p<sup>au<\/sup><\/td>\n<td><\/td>\n<td style=\"text-align: center;\">p<sup>au<\/sup><\/td>\n<td style=\"text-align: center;\">p<sup>au<\/sup><\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\"><\/th>\n<td style=\"text-align: center;\">P<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<td style=\"text-align: center;\">p<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<td><\/td>\n<td style=\"text-align: center;\">P<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<td style=\"text-align: center;\">p<sup>au<\/sup>p<sup>au<\/sup><\/td>\n<\/tr>\n<\/tbody>\n<tfoot>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\" colspan=\"2\">2 (0,5P \u00d7 0,5p) = 50%<\/td>\n<td><\/td>\n<td style=\"text-align: center;\" colspan=\"2\">verde \/ arlequim recessivo<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\" colspan=\"2\">2 (0,5p \u00d7 0,5p) = 50%<\/td>\n<td><\/td>\n<td style=\"text-align: center;\" colspan=\"2\">arlequim recessivo<\/td>\n<\/tr>\n<\/tfoot>\n<\/table>\n<h3>2\u00aa Lei de Mendel<\/h3>\n<p>A primeira lei de Mendel, embora seja aplic\u00e1vel nos Agapornis, torna-se restrita pois ao ser elaborada n\u00e3o considerava a intera\u00e7\u00e3o g\u00eanica.<\/p>\n<p>O esquizocromismo faz com que a totalidade do car\u00e1ter gen\u00e9tico para cor nos Agapornis sofram intera\u00e7\u00e3o g\u00eanica.O que se mais se aplica no agapornismo \u00e9 a 2\u00aa Lei de Mendel que analisa dois ou mais pares de genes ao mesmo tempo. (D\u2019Angieri, A. \u2013 \u201cThe Colored Atlas of Lovebirds\u201d \u2013 TFH \u2013 1997)<\/p>\n<p>A segrega\u00e7\u00e3o, neste caso, ocorre independentemente para cada par de genes, ou seja: cada indiv\u00edduo produzir\u00e1 gametas portadores de apenas um gene de cada par, de acordo com a 1\u00aa Lei de Mendel.<\/p>\n<p>&nbsp;<\/p>\n<table>\n<caption>Exemplo 4<\/caption>\n<thead>\n<tr>\n<th style=\"text-align: center;\"><\/th>\n<th style=\"text-align: center;\" colspan=\"2\">golden americano (g<sup>a<\/sup>g<sup>a<\/sup>)<\/th>\n<th>\u00d7<\/th>\n<th style=\"text-align: center;\" colspan=\"2\">aqua (pp)<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\" colspan=\"5\">Obs: Os genes \u201cG<sup>a<\/sup>\u201d e \u201cP\u201d indicam aus\u00eancia dos fatores amarelos (g<sup>a<\/sup>) e aqua (p) respectivamente.<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\" colspan=\"2\">g<sup>a<\/sup>g<sup>a<\/sup>PP<\/td>\n<td style=\"text-align: center;\">\u00d7<\/td>\n<td style=\"text-align: center;\" colspan=\"2\">G<sup>a<\/sup>G<sup>a<\/sup>pp<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\">50%<\/td>\n<td style=\"text-align: center;\">50%<\/td>\n<td><\/td>\n<td style=\"text-align: center;\">50%<\/td>\n<td style=\"text-align: center;\">50%<\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">G:<\/th>\n<td style=\"text-align: center;\">g<sup>a<\/sup>P<\/td>\n<td style=\"text-align: center;\">g<sup>a<\/sup>P<\/td>\n<td><\/td>\n<td style=\"text-align: center;\">G<sup>a<\/sup>p<\/td>\n<td style=\"text-align: center;\">G<sup>a<\/sup>p<\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">F1:<\/th>\n<td style=\"text-align: center;\" colspan=\"5\">G<sup>a<\/sup>g<sup>a<\/sup>Pp<\/td>\n<\/tr>\n<\/tbody>\n<tfoot>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\" colspan=\"5\">100% verde<\/td>\n<\/tr>\n<\/tfoot>\n<\/table>\n<p>O duplo heterozigoto G<sup>a<\/sup>g<sup>a<\/sup>Pp possui o fen\u00f3tipo verde, uma vez que ambos os genes s\u00e3o recessivos e est\u00e3o em heterozigose: temos um indiv\u00edduo N\u00c3O GOLDEN (G<sup>a<\/sup>) e N\u00c3O AQUA (P), isto \u00e9, um indiv\u00edduo que n\u00e3o mostra nenhum fen\u00f3tipo mutante vis\u00edvel s\u00f3 pode mostrar o fen\u00f3tipo selvagem n\u00e3o mutante (verde).<\/p>\n<table>\n<caption>F2:<\/caption>\n<thead>\n<tr>\n<th colspan=\"4\">G<sup>a<\/sup>aPp<\/th>\n<th>\u00d7<\/th>\n<th colspan=\"4\">G<sup>a<\/sup>g<sup>a<\/sup>Pp<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td>25%<\/td>\n<td>25%<\/td>\n<td>25%<\/td>\n<td>25%<\/td>\n<td><\/td>\n<td>25%<\/td>\n<td>25%<\/td>\n<td>25%<\/td>\n<td>25%<\/td>\n<\/tr>\n<tr>\n<td>G<sup>a<\/sup>p<\/td>\n<td>G<sup>a<\/sup>p<\/td>\n<td>g<sup>a<\/sup>P<\/td>\n<td>g<sup>a<\/sup>p<\/td>\n<td><\/td>\n<td>G<sup>a<\/sup>P<\/td>\n<td>G<sup>a<\/sup>p<\/td>\n<td>g<sup>a<\/sup>P<\/td>\n<td>g<sup>a<\/sup>p<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<table>\n<caption>F2:<\/caption>\n<thead>\n<tr>\n<th><\/th>\n<th>G<sup>a<\/sup>P<\/th>\n<th>G<sup>a<\/sup>p<\/th>\n<th>g<sup>a<\/sup>P<\/th>\n<th>g<sup>a<\/sup>p<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<th scope=\"row\">G<sup>a<\/sup>P<\/th>\n<td>G<sup>a<\/sup>G<sup>a<\/sup>PP<\/td>\n<td>G<sup>a<\/sup>G<sup>a<\/sup>Pp<\/td>\n<td>G<sup>a<\/sup>g<sup>a<\/sup>PP<\/td>\n<td>G<sup>a<\/sup>g<sup>a<\/sup>pp<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">G<sup>a<\/sup>p<\/th>\n<td>G<sup>a<\/sup>g<sup>a<\/sup>Pp<\/td>\n<td>G<sup>a<\/sup>G<sup>a<\/sup>pp<\/td>\n<td>G<sup>a<\/sup>g<sup>a<\/sup>Pp<\/td>\n<td>G<sup>a<\/sup>g<sup>a<\/sup>pp<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">g<sup>a<\/sup>P<\/th>\n<td>G<sup>a<\/sup>g<sup>a<\/sup>PP<\/td>\n<td>G<sup>a<\/sup>g<sup>a<\/sup>Pp<\/td>\n<td>g<sup>a<\/sup>g<sup>a<\/sup>PP<\/td>\n<td>g<sup>a<\/sup>g<sup>a<\/sup>Pp<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">g<sup>a<\/sup>p<\/th>\n<td>G<sup>a<\/sup>g<sup>a<\/sup>Pp<\/td>\n<td>G<sup>a<\/sup>g<sup>a<\/sup>pp<\/td>\n<td>g<sup>a<\/sup>g<sup>a<\/sup>Pp<\/td>\n<td>g<sup>a<\/sup>g<sup>a<\/sup>pp<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>Cada quadrinho corresponde a uma freq\u00fc\u00eancia de 1\/16 = 6,25%.<\/p>\n<table>\n<thead>\n<tr>\n<th>Gen\u00f3tipo<\/th>\n<th>freq\u00fc\u00eancia (%)<\/th>\n<th>fen\u00f3tipo<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td>G<sup>a<\/sup>G<sup>a<\/sup>PP<\/td>\n<td>6,25<\/td>\n<td>verde<\/td>\n<\/tr>\n<tr>\n<td>G<sup>a<\/sup>G<sup>a<\/sup>Pp<\/td>\n<td>12,50<\/td>\n<td>verde\/aqua<\/td>\n<\/tr>\n<tr>\n<td>G<sup>a<\/sup>g<sup>a<\/sup>PP<\/td>\n<td>12,50<\/td>\n<td>verde\/golden americano<\/td>\n<\/tr>\n<tr>\n<td>G<sup>a<\/sup>g<sup>a<\/sup>Pp<\/td>\n<td>25,00<\/td>\n<td>verde\/golden, americano e aqua<\/td>\n<\/tr>\n<tr>\n<td>G<sup>a<\/sup>G<sup>a<\/sup>pp<\/td>\n<td>6,25<\/td>\n<td>aqua<\/td>\n<\/tr>\n<tr>\n<td>G<sup>a<\/sup>g<sup>a<\/sup>pp<\/td>\n<td>12,50<\/td>\n<td>aqua\/golden americano<\/td>\n<\/tr>\n<tr>\n<td>g<sup>a<\/sup>g<sup>a<\/sup>Pp<\/td>\n<td>12,50<\/td>\n<td>golden.amer. \/ aqua e prata<\/td>\n<\/tr>\n<tr>\n<td>g<sup>a<\/sup>g<sup>a<\/sup>PP<\/td>\n<td>6,25<\/td>\n<td>golden americano<\/td>\n<\/tr>\n<tr>\n<td>g<sup>a<\/sup>g<sup>a<\/sup>pp<\/td>\n<td>6,25<\/td>\n<td>prata (aqua escamado dilu\u00eddo)<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>Nota: a intera\u00e7\u00e3o dos genes \u201cp\u201d e \u201cg<sup>a<\/sup>\u201d se mostra no duplo homozigoto \u201cg<sup>a<\/sup>g<sup>a<\/sup>pp\u201d, que possui a cor (fen\u00f3tipo) prata, uma combina\u00e7\u00e3o inexistente at\u00e9 ent\u00e3o. Este resultado mostra quatro cores (fen\u00f3tipos) diferentes: verde, amarelo (golden) americano, aqua e prata (silver americano ou aqua escamado dilu\u00eddo).<\/p>\n<p>O exemplo acima \u00e9 t\u00edpico do funcionamento da maioria dos genes dos Agapornis, demonstrando claramente os fen\u00f4menos de domin\u00e2ncia, recessividade e intera\u00e7\u00e3o g\u00eanica (os fatores aqua e amarelo americano se interagem formando uma quarta cor: o prata ou silver hoje tamb\u00e9m conhecido como aqua escamado dilu\u00eddo.<\/p>\n<table>\n<caption>Exemplo 5<\/caption>\n<thead>\n<tr>\n<th style=\"text-align: center;\"><\/th>\n<th style=\"text-align: center;\" colspan=\"2\">aqua \/ escamado (golden)<\/th>\n<th>\u00d7<\/th>\n<th style=\"text-align: center;\" colspan=\"2\">aqua \/ escamado (golden)<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td style=\"text-align: center;\"><\/td>\n<td style=\"text-align: center;\" colspan=\"2\">G<sup>a<\/sup>g<sup>a<\/sup>pp<\/td>\n<td>\u00d7<\/td>\n<td style=\"text-align: center;\" colspan=\"2\">G<sup>a<\/sup>g<sup>a<\/sup>pp<\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">G:<\/th>\n<td style=\"text-align: center;\">G<sup>a<\/sup>p<\/td>\n<td style=\"text-align: center;\">g<sup>a<\/sup>p<\/td>\n<td><\/td>\n<td style=\"text-align: center;\">G<sup>a<\/sup>p<\/td>\n<td style=\"text-align: center;\">g<sup>a<\/sup>p<\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">F1:<\/th>\n<td style=\"text-align: center;\">G<sup>a<\/sup>G<sup>a<\/sup>pp<\/td>\n<td style=\"text-align: center;\" colspan=\"3\">G<sup>a<\/sup>g<sup>a<\/sup>pp<\/td>\n<td style=\"text-align: center;\">g<sup>a<\/sup>g<sup>a<\/sup>pp<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>Nota: neste caso, embora seja dois os pares de genes envolvidos, ambos os pais s\u00e3o homozigotos para aqua (p), assim as propor\u00e7\u00f5es obtidas seguem a 1\u00aa Lei de Mendel uma vez que n\u00e3o h\u00e1 variabilidade no locus do fator aqua.<\/p>\n<table>\n<caption>Exemplo 6<\/caption>\n<thead>\n<tr>\n<th style=\"text-align: center;\"><\/th>\n<th style=\"text-align: center;\" colspan=\"4\">personatus azul<\/th>\n<th>\u00d7<\/th>\n<th style=\"text-align: center;\" colspan=\"4\">personatus verde dilu\u00eddo (verde pastel)<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\" colspan=\"4\">aaII<\/td>\n<td>\u00d7<\/td>\n<td style=\"text-align: center;\" colspan=\"4\">AAi<sup>d<\/sup>i<sup>d<\/sup><\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">G:<\/th>\n<td style=\"text-align: center;\" colspan=\"2\">aI<\/td>\n<td style=\"text-align: center;\" colspan=\"2\">aI<\/td>\n<td><\/td>\n<td style=\"text-align: center;\" colspan=\"2\">Ai<sup>d<\/sup><\/td>\n<td style=\"text-align: center;\" colspan=\"2\">Ai<sup>d<\/sup><\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">F1:<\/th>\n<td style=\"text-align: center;\" colspan=\"9\">AaIi<sup>d<\/sup><\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td colspan=\"9\">100% personatus verde \/ azul e pastel<\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">F2:<\/th>\n<td style=\"text-align: center;\" colspan=\"4\">AaIi<sup>d<\/sup><\/td>\n<td>\u00d7<\/td>\n<td style=\"text-align: center;\" colspan=\"4\">AaIi<sup>d<\/sup><\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">G:<\/th>\n<td style=\"text-align: center;\">AI<\/td>\n<td style=\"text-align: center;\">Ai<sup>d<\/sup><\/td>\n<td style=\"text-align: center;\">aI<\/td>\n<td style=\"text-align: center;\">ai<sup>d<\/sup><\/td>\n<td><\/td>\n<td style=\"text-align: center;\">AI<\/td>\n<td style=\"text-align: center;\">Ai<sup>d<\/sup><\/td>\n<td style=\"text-align: center;\">aI<\/td>\n<td style=\"text-align: center;\">ai<sup>d<\/sup><\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<table>\n<thead>\n<tr>\n<th><\/th>\n<th>AI<\/th>\n<th>Ai<sup>d<\/sup><\/th>\n<th>aI<\/th>\n<th>ai<sup>d<\/sup><\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<th scope=\"row\">AI<\/th>\n<td>AAII<\/td>\n<td>AAId<\/td>\n<td>AaII<\/td>\n<td>AaIi<sup>d<\/sup><\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">Ai<sup>d<\/sup><\/th>\n<td>AAIi<sup>d<\/sup><\/td>\n<td>AAi<sup>d<\/sup>i<sup>d<\/sup><\/td>\n<td>AaIi<sup>d<\/sup><\/td>\n<td>Aai<sup>d<\/sup>i<sup>d<\/sup><\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">aI<\/th>\n<td>AaII<\/td>\n<td>AaIi<sup>d<\/sup><\/td>\n<td>aaII<\/td>\n<td>aaIi<sup>d<\/sup><\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">ai<sup>d<\/sup><\/th>\n<td>AaIi<sup>d<\/sup><\/td>\n<td>Aai<sup>d<\/sup>i<sup>d<\/sup><\/td>\n<td>aaIi<sup>d<\/sup><\/td>\n<td>aai<sup>d<\/sup>i<sup>d<\/sup><\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<table>\n<tbody>\n<tr>\n<th scope=\"row\">AAII:<\/th>\n<td>verde<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">AAIi<sup>d<\/sup>:<\/th>\n<td>verde \/ pastel (dilu\u00eddo)<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">Aai<sup>d<\/sup>i<sup>d<\/sup><\/th>\n<td>verde pastel (dilu\u00eddo) \/ azul<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">AaIi<sup>d<\/sup>:<\/th>\n<td>verde \/ azul e pastel (dilu\u00eddo)<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">AaII:<\/th>\n<td>verde \/ azul<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">AAi<sup>d<\/sup>i<sup>d<\/sup>:<\/th>\n<td>verde pastel (dilu\u00eddo)<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">aaIi<sup>d<\/sup>:<\/th>\n<td>azul \/ pastel (dilu\u00eddo)<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">aaII:<\/th>\n<td>azul<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">aai<sup>d<\/sup>i<sup>d<\/sup>:<\/th>\n<td>azul pastel (dilu\u00eddo)<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>Podemos e devemos avaliar tr\u00eas pares de genes ao mesmo tempo:<\/p>\n<table>\n<caption>Exemplo 7<\/caption>\n<thead>\n<tr>\n<th style=\"text-align: center;\"><\/th>\n<th style=\"text-align: center;\" colspan=\"4\">personatus azul pastel (dilu\u00eddo)<\/th>\n<th>\u00d7<\/th>\n<th style=\"text-align: center;\" colspan=\"4\">personatus oliva<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<th scope=\"row\"><\/th>\n<td style=\"text-align: center;\" colspan=\"4\">i<sup>d<\/sup>i<sup>d<\/sup>aadd<\/td>\n<td>\u00d7<\/td>\n<td style=\"text-align: center;\" colspan=\"4\">IIAADD<\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">G:<\/th>\n<td style=\"text-align: center;\" colspan=\"4\">i<sup>d<\/sup>ad<\/td>\n<td><\/td>\n<td style=\"text-align: center;\" colspan=\"4\">IAD<\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">F1:<\/th>\n<td style=\"text-align: center;\" colspan=\"9\">Ii<sup>d<\/sup>AaDd<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td colspan=\"9\">100% verdes jade<\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">F2:<\/th>\n<td style=\"text-align: center;\" colspan=\"4\">Ii<sup>d<\/sup>AaDd<\/td>\n<td>\u00d7<\/td>\n<td style=\"text-align: center;\" colspan=\"4\">Ii<sup>d<\/sup>AaDd<\/td>\n<\/tr>\n<tr>\n<th style=\"text-align: left;\" scope=\"row\">G:<\/th>\n<td style=\"text-align: center;\" colspan=\"4\">I<\/td>\n<td><\/td>\n<td style=\"text-align: center;\" colspan=\"4\">i<sup>d<\/sup><\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td colspan=\"4\">50%<\/td>\n<td><\/td>\n<td colspan=\"4\">50%<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td colspan=\"2\">a<\/td>\n<td colspan=\"2\">A<\/td>\n<td><\/td>\n<td colspan=\"2\">a<\/td>\n<td colspan=\"2\">A<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td colspan=\"2\">50%<\/td>\n<td colspan=\"2\">50%<\/td>\n<td><\/td>\n<td colspan=\"2\">50%<\/td>\n<td colspan=\"2\">50%<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td>d<\/td>\n<td>D<\/td>\n<td>d<\/td>\n<td>D<\/td>\n<td><\/td>\n<td>d<\/td>\n<td>D<\/td>\n<td>d<\/td>\n<td>D<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td>50%<\/td>\n<td>50%<\/td>\n<td>50%<\/td>\n<td>50%<\/td>\n<td><\/td>\n<td>50%<\/td>\n<td>50%<\/td>\n<td>50%<\/td>\n<td>50%<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td>Iad<\/td>\n<td>IaD<\/td>\n<td>IAd<\/td>\n<td>IAD<\/td>\n<td><\/td>\n<td>i<sup>d<\/sup>ad<\/td>\n<td>i<sup>d<\/sup>aD<\/td>\n<td>i<sup>d<\/sup>Ad<\/td>\n<td>i<sup>d<\/sup>AD<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\"><\/th>\n<td>12,5%<\/td>\n<td>12,5%<\/td>\n<td>12,5%<\/td>\n<td>12,5%<\/td>\n<td><\/td>\n<td>12,5%<\/td>\n<td>12,5%<\/td>\n<td>12,5%<\/td>\n<td>12,5%<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<table>\n<thead>\n<tr>\n<th><\/th>\n<th><\/th>\n<th>IAD<\/th>\n<th>IAd<\/th>\n<th>Iad<\/th>\n<th>i<sup>d<\/sup>AD<\/th>\n<th>i<sup>d<\/sup>Ad<\/th>\n<th>i<sup>d<\/sup>aD<\/th>\n<th>i<sup>d<\/sup>ad<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<th scope=\"row\">IAd<\/th>\n<td>IIAADd<\/td>\n<td>IIAAdd<\/td>\n<td>IIAaDd<\/td>\n<td>IIAadd<\/td>\n<td>Ii<sup>d<\/sup>AADd<\/td>\n<td>Ii<sup>d<\/sup>AAdd<\/td>\n<td>Ii<sup>d<\/sup>AaDd<\/td>\n<td>Ii<sup>d<\/sup>Aadd<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">IaD<\/th>\n<td>IIAaDd<\/td>\n<td>IIAaDd<\/td>\n<td>IIaaDD<\/td>\n<td>IIaaDd<\/td>\n<td>Ii<sup>d<\/sup>AaDD<\/td>\n<td>Ii<sup>d<\/sup>AaDd<\/td>\n<td>Ii<sup>d<\/sup>aaDD<\/td>\n<td>Ii<sup>d<\/sup>aaDd<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">Iad<\/th>\n<td>IIaADd<\/td>\n<td>IIAadd<\/td>\n<td>IIaaDd<\/td>\n<td>IIaadd<\/td>\n<td>Ii<sup>d<\/sup>AaDd<\/td>\n<td>Ii<sup>d<\/sup>Aadd<\/td>\n<td>Ii<sup>d<\/sup>aaDd<\/td>\n<td>Ii<sup>d<\/sup>aadd<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">i<sup>d<\/sup>AD<\/th>\n<td>Ii<sup>d<\/sup>AADD<\/td>\n<td>Ii<sup>d<\/sup>AADd<\/td>\n<td>IiAaDD<\/td>\n<td>Ii<sup>d<\/sup>AaDd<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>AADD<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>AADd<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>AaDD<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>AaDd<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">i<sup>d<\/sup>Ad<\/th>\n<td>Ii<sup>d<\/sup>AADd<\/td>\n<td>Ii<sup>d<\/sup>AAdd<\/td>\n<td>Ii<sup>d<\/sup>Aadd<\/td>\n<td>Ii<sup>d<\/sup>Aadd<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>AADd<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>Aadd<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>AaDd<\/td>\n<td>iiAadd<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">i<sup>d<\/sup>aD<\/th>\n<td>Ii<sup>d<\/sup>AaDD<\/td>\n<td>Ii<sup>d<\/sup>AaDd<\/td>\n<td>Ii<sup>d<\/sup>aaDD<\/td>\n<td>Ii<sup>d<\/sup>aaDd<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>AaDD<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>AaDd<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>aaDD<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>aaDd<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">i<sup>d<\/sup>ad<\/th>\n<td>Ii<sup>d<\/sup>AaDd<\/td>\n<td>Ii<sup>d<\/sup>Aadd<\/td>\n<td>Ii<sup>d<\/sup>aaDd<\/td>\n<td>Ii<sup>d<\/sup>aadd<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>AaDd<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>Aadd<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>aaDd<\/td>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>aadd<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>Teremos filhotes de doze cores (fen\u00f3tipos) diferentes:<\/p>\n<table>\n<thead>\n<tr>\n<th>fen\u00f3tipo<\/th>\n<th>gen\u00f3tipo<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<th scope=\"row\">verde jade<\/th>\n<td>(D)I_ A_D\u00ad_A\u00ad_dd<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">verde<\/th>\n<td>I\u00ad_ A_ dd<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">oliva<\/th>\n<td>I_ AaDD<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">oliva pastel (dilu\u00eddo)<\/th>\n<td>i<sup>d<\/sup>i<sup>d <\/sup>A _ DD<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">verde pastel (dilu\u00eddo)<\/th>\n<td>i<sup>d<\/sup>i<sup><sup>d<\/sup><\/sup><\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">verde jade pastel (dilu\u00eddo)<\/th>\n<td>i<sup>d<\/sup>i<sup>d<\/sup> A _ DD<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">azul cobalto<\/th>\n<td>I_ aa D\u00ad_<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">azul<\/th>\n<td>I_ aadd<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">azul pastel (dilu\u00eddo)<\/th>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>aadd<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">azul cobalto pastel (dilu\u00eddo)<\/th>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>iaaD_<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">malva<\/th>\n<td>I_aaDD<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">malva pastel (dilu\u00eddo)<\/th>\n<td>i<sup>d<\/sup>i<sup>d<\/sup>aaDD<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>( _ ): significa que n\u00e3o importa o gene presente pois n\u00e3o haver\u00e1 mudan\u00e7a no fen\u00f3tipo.<\/p>\n<h2>Vincula\u00e7\u00e3o (linkage) e Permuta<\/h2>\n<p>A lei de segrega\u00e7\u00e3o dos gametas ( 2\u00ba lei de Mendel ), que acabamos de ver, n\u00e3o \u00e9 aplic\u00e1vel a todos os casos, s\u00f3 ocorre entre genes situados em cromossomos n\u00e3o hom\u00f3logos.<br \/>\nOs genes se situam linearmente nos cromossomos e deveriam originar, pela l\u00f3gica, dois outros indiv\u00edduos iguais quando ocorresse a duplica\u00e7\u00e3o g\u00eanica. A pr\u00e1tica nos mostra que ainda assim a recombina\u00e7\u00e3o g\u00eanica ocorre.<br \/>\nEm um mesmo cromossomos os genes est\u00e3o \u201cligados\u201d ou \u201cvinculados\u201d (linked) e tendem a permanecer unidos, fen\u00f4meno este que foi definido como \u201cVincula\u00e7\u00e3o\u201d ou \u201cLinkage\u201d.<br \/>\nA recombina\u00e7\u00e3o dos genes em \u201clinkage\u201d se d\u00e1 atrav\u00e9s de um outro fen\u00f4meno: o \u201ccrossing-over\u201d.<br \/>\nO \u201ccrossing-over\u201d \u00e9 uma permuta que se d\u00e1 ocasionalmente, sem nenhuma ordem ou regra preestabelecida, em qualquer ponto dos filamentos ( crom\u00e1tides ) dos cromossomos hom\u00f3logos, numa t\u00e9trade ( conjunto de quatro crom\u00e1tides na duplica\u00e7\u00e3o dos cromossomos hom\u00f3logos ). Ap\u00f3s um entrela\u00e7amento os filamentos se repelem, havendo uma permuta de fragmentos onde se situam diversos genes. Tal mecanismo substitui apenas em parte a impossibilidade de recombina\u00e7\u00e3o pela 2\u00ba lei de Mendel.<br \/>\nA \u201ctaxa de permuta\u201d entre dois genes depende e \u00e9 diretamente proporcional a dist\u00e2ncia que os separa nos cromossomos: quanto mais distantes maior \u00e9 a taxa de crossing e vice-versa, num m\u00e1ximo de 50%.<br \/>\nO fen\u00f4meno \u201clinkage\u201d nos Agapornis, juntamente com o crossing-over, ocorre de maneira clara no fator escuro (vide fator escurecedor ) e entre este e o fator aqua.<br \/>\nUtilizemos o fator escurecedor (D) e fator aqua (p); se houver \u201clinkage\u201d completo entre eles, cada gameta dever\u00e1 levar D-P ou d-p e n\u00e3o ocorrer\u00e1 recombina\u00e7\u00e3o, havendo somente dois tipos de gametas DP (50%) e d-p (50%). Contudo sabemos que existem indiv\u00edduos malvas (D-p) e que nos indica que ocorreu a permuta.<\/p>\n<table>\n<caption>Exemplo 8: roseicollis<\/caption>\n<thead>\n<tr>\n<th><\/th>\n<th colspan=\"4\">oliva<\/th>\n<th>\u00d7<\/th>\n<th colspan=\"4\">aqua<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td><\/td>\n<td colspan=\"4\">DDPP<\/td>\n<td><\/td>\n<td colspan=\"4\">ddpp<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">F1:<\/th>\n<td colspan=\"4\">DdPp<\/td>\n<td><\/td>\n<td colspan=\"4\">100% jades<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">F2:<\/th>\n<td colspan=\"4\">DdPp<\/td>\n<th>\u00d7<\/th>\n<td colspan=\"4\">DdPp<\/td>\n<\/tr>\n<tr>\n<td><\/td>\n<td>DP<\/td>\n<td>Dp<\/td>\n<td>dP<\/td>\n<td>dp<\/td>\n<td><\/td>\n<td>DP<\/td>\n<td>Dp<\/td>\n<td>dP<\/td>\n<td>dp<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<table>\n<thead>\n<tr>\n<th><\/th>\n<th>DP<\/th>\n<th>Dp<\/th>\n<th>dP<\/th>\n<th>dp<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<th scope=\"row\">DP<\/th>\n<td>DDPP<\/td>\n<td>DDPp<\/td>\n<td>DdPP<\/td>\n<td>DdPp<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">Dp<\/th>\n<td>DDPp<\/td>\n<td>DDpp<\/td>\n<td>DdPp<\/td>\n<td>Ddpp<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">dP<\/th>\n<td>DdPP<\/td>\n<td>DdPp<\/td>\n<td>ddPP<\/td>\n<td>ddPp<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">dP<\/th>\n<td>DdPp<\/td>\n<td>Ddpp<\/td>\n<td>ddPp<\/td>\n<td>ddpp<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>Teoricamente, dever\u00edamos ter malvas (DDpp) \u00e0 1\/16 = 6,25%. O observado foi que as malvas no cruzamento acima surgem em freq\u00fc\u00eancia de 1\/93 ou seja 1,07% (D\u2019Angieri \u2013 \u201cThe colored Atlas of Lovebirds\u201d) . Ou seja 98,93 % das vezes n\u00e3o vai ocorrer a troca de genes.<\/p>\n<p>Linkage: os genes pr\u00f3ximos n\u00e3o permitem a permuta, fato que ocorre na realidade entre os fatores vistos, reduzindo assim de 6,25% (esperado) para 1,07% (obtido) o percentual de malvas no cruzamento acima.<\/p>\n<p>Identificamos assim dois tipos de heterozigotos para fator escurecedor e pastel:<\/p>\n<table>\n<thead>\n<tr>\n<th colspan=\"2\">tipo I<\/th>\n<th colspan=\"2\">tipo II<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td>D<\/td>\n<td>d<\/td>\n<td>D<\/td>\n<td>d<\/td>\n<\/tr>\n<tr>\n<td>P<\/td>\n<td>p<\/td>\n<td>p<\/td>\n<td>P<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>Ambos os indiv\u00edduos acima s\u00e3o jades, por\u00e9m apenas o tipo II produzir\u00e1 malvas (DDpp) na freq\u00fc\u00eancia mendeliana.<br \/>\nSendo assim faz-se necess\u00e1rio primeiro obtermos os jades tipo II de cruzamentos com o tipo I, para depois conseguirmos manter malvas em grandes<br \/>\nfrequ\u00eancia.<\/p>\n<p>A m\u00e1xima taxa de crossing ou taxa de permuta \u00e9 50% o que significa uma permuta, em 100% das c\u00e9lulas, entre dois genes.<\/p>\n<h3>Genes Vinculados ao Sexo (Sex-Linkage)<\/h3>\n<p>Existem diversos genes que n\u00e3o se manifestam usualmente e em alguns cruzamentos podemos notar que determinados fen\u00f3tipos aparecem predominantemente em f\u00eameas ou machos.<\/p>\n<p>A ocorr\u00eancia de genes vinculados ao sexo \u00e9 conhecida como \u201csex-linkage\u201d. Estes genes est\u00e3o localizados nos cromossomos sexuais ou heterocromossomos \u201cZ\u201d e \u201cW\u201d no caso das aves. O uso de Z e W significa automaticamente que o sexo heterogam\u00e9tico \u00e9 o feminino ( ZW ):<\/p>\n<p>Os cromossomos Z e W s\u00e3o, respectivamente, an\u00e1logos a X e Y de outras classes animais (mam\u00edferos, alguns insetos, etc.).Infelizmente a grande maioria dos autores n\u00e3o cient\u00edficos de Agapornis e Aves em geral se utilizam X e Y quando aborda muta\u00e7\u00f5es ligadas ao sexo. Embora sejam s\u00f3 s\u00edmbolos devemos respeitar a nomenclatura cient\u00edfica gen\u00e9tica geral (D\u2019Angieri, A., The Colored Atlas of Lovebirds-TFH -1997).<\/p>\n<p>\u201cO uso de XY e XX para aves n\u00e3o \u00e9 adequado, pois uma vez utilizados, assume-se que o modelo de sexo heterogam\u00e9tico \u00e9 representado pelos machos e isto n\u00e3o ocorre nas aves\u201d (D\u2019Angieri,A, The Colored Atlas of Lovebirds-TFH -1997)<\/p>\n<blockquote><p>\u201cNas aves \u00e9 a f\u00eamea que determina o sexo dos filhotes atrav\u00e9s do cromossomo W.\u201d (Buckley, 1987).<\/p><\/blockquote>\n<p>Um gene ligado ao sexo, se recessivo, s\u00f3 se expressar\u00e1 nas aves, quando em homozigose ( presente em ambos Zs ); nas f\u00eameas hemezig\u00f3ticas apenas com um gene obteremos o fen\u00f3tipo em quest\u00e3o. Assim, os caracteres sex-linked recessivos aparecem mais freq\u00fcentemente nas f\u00eameas, levando a uma popula\u00e7\u00e3o feminina maior que a masculina, fato facilmente observado nos fatores ino , opalino, p\u00e1lido e canela dos roseicollis.<\/p>\n<p>Devemos ainda lembrar que as aves possuem um cari\u00f3tipo (mapa cromoss\u00f4mico) uma cadeia dipl\u00f3ide 40-80, portanto casos de linkage s\u00e3o regras e n\u00e3o exce\u00e7\u00e3o (Buckley, 1987).<\/p>\n<p>&nbsp;<\/p>\n<table>\n<caption>Exemplo 9<\/caption>\n<thead>\n<tr>\n<th><\/th>\n<th colspan=\"2\">opalino verde<\/th>\n<th>\u00d7<\/th>\n<th colspan=\"2\">aqua<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td><\/td>\n<td colspan=\"2\">Z<sup>o<\/sup>Z<sup>o<\/sup>PP<\/td>\n<td><\/td>\n<td colspan=\"2\">Z<sup>O<\/sup>Wpp<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">G:<\/th>\n<td>Z<sup>o<\/sup>P<\/td>\n<td>Z<sup>o<\/sup>P<\/td>\n<td><\/td>\n<td>Z<sup>O<\/sup>p<\/td>\n<td>Wp<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">F1:<\/th>\n<td colspan=\"2\">Z<sup>O<\/sup>Z<sup>o<\/sup>Pp<\/td>\n<td><\/td>\n<td colspan=\"2\">Z<sup>o<\/sup>WPp<\/td>\n<\/tr>\n<\/tbody>\n<tfoot>\n<tr>\n<td><\/td>\n<td colspan=\"2\">100% dos machos<br \/>\nverdes\/opalino e aqua<\/td>\n<td><\/td>\n<td colspan=\"2\">100% das f\u00eameas<br \/>\nopalinas verdes \/ aqua<\/td>\n<\/tr>\n<\/tfoot>\n<\/table>\n<table>\n<caption>Exemplo 10<\/caption>\n<thead>\n<tr>\n<th><\/th>\n<th colspan=\"4\">verde\/opalino e aqua<\/th>\n<th>\u00d7<\/th>\n<th colspan=\"4\">aqua<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td><\/td>\n<td colspan=\"4\">Z<sup>o<\/sup>Z<sup>o<\/sup>Pp<\/td>\n<td><\/td>\n<td colspan=\"4\">Z<sup>o<\/sup>Wpp<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">G:<\/th>\n<td>Z<sup>o<\/sup>P<\/td>\n<td>Z<sup>o<\/sup>p<\/td>\n<td>Z<sup>o<\/sup>p<\/td>\n<td>Z<sup>o<\/sup>P<\/td>\n<td><\/td>\n<td colspan=\"2\">Z<sup>o<\/sup>p<\/td>\n<td colspan=\"2\">Wp<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<table>\n<tbody>\n<tr>\n<th scope=\"row\">F1:<\/th>\n<td>Z<sup>o<\/sup>P<\/td>\n<td>Z<sup>o<\/sup>p<\/td>\n<td>Z<sup>o<\/sup>P<\/td>\n<td>Z<sup>o<\/sup>p<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">Z<sup>o<\/sup>p<\/th>\n<td>Z<sup>o<\/sup>Z<sup>o<\/sup>Pp<\/td>\n<td>Z<sup>o<\/sup>Z<sup>o<\/sup>pp<\/td>\n<td>Z<sup>o<\/sup>Z<sup>i<\/sup>Pp<\/td>\n<td>Z<sup>o<\/sup>Z<sup>o<\/sup>pp<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">Wp<\/th>\n<td>Z<sup>o<\/sup>WPp<\/td>\n<td>Z<sup>o<\/sup>Wpp<\/td>\n<td>Z<sup>o<\/sup>WPp<\/td>\n<td>Z<sup>o<\/sup>Wpp<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<table>\n<tbody>\n<tr>\n<th rowspan=\"2\" scope=\"row\">machos:<\/th>\n<td>Z<sup>o<\/sup>Z<sup>o<\/sup>Pp<\/td>\n<td>verde \/ opalino, aqua<\/td>\n<td>25%<\/td>\n<\/tr>\n<tr>\n<td>Z<sup>o<\/sup>Z<sup>o<\/sup>pp<\/td>\n<td>aqua \/ opalino<\/td>\n<td>25%<\/td>\n<\/tr>\n<tr>\n<th rowspan=\"4\" scope=\"row\">f\u00eameas:<\/th>\n<td>Z<sup>o<\/sup>WPp<\/td>\n<td>opalino<\/td>\n<td>12,5%<\/td>\n<\/tr>\n<tr>\n<td>Z<sup>I<\/sup>Wpp<\/td>\n<td>aqua<\/td>\n<td>12,5%<\/td>\n<\/tr>\n<tr>\n<td>Z<sup>I<\/sup>WPp<\/td>\n<td>verde<\/td>\n<td>12,5%<\/td>\n<\/tr>\n<tr>\n<td>Z<sup>i<\/sup>Wpp<\/td>\n<td>opalino aqua<\/td>\n<td>12,5%<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<h3>Alelos M\u00faltiplos<\/h3>\n<p>A polialelia (alelos m\u00faltiplos) \u00e9 um caso peculiar de gen\u00e9tica e que se faz representar nos Agapornis.<\/p>\n<p>Este fen\u00f4meno consiste na exist\u00eancia de mais de um gene alelo para um mesmo locus e conseq\u00fcentemente um mesmo car\u00e1ter. Estes genes s\u00e3o variantes de muta\u00e7\u00f5es que ocorreram no mesmo gene em ocasi\u00f5es diferentes.<\/p>\n<p>O fator ino dos roseicollis \u00e9 um exemplo t\u00edpico de alelos m\u00faltiplos: duas muta\u00e7\u00f5es ocorreram separadamente em dois locais distantes do globo \u2013 um na Austr\u00e1lia em 1957 e outro nos Estados Unidos em 1973.<\/p>\n<p>A muta\u00e7\u00e3o australiana, erroneamente conhecida como \u201ccanela\u201d australiano \u00e9, na verdade um caso de esquizocromia parcial ( lutinismo ) enquanto que seu alelo americano \u00e9 um caso cl\u00e1ssico de esquizocr\u00f4mico lutino. Ambos os genes em quest\u00e3o ocupam o mesmo \u201clocus\u201d e o alelo australiano \u00e9 dominante em rela\u00e7\u00e3o ao americano. Separadamente s\u00e3o ambos recessivos (vide fator ino). (D\u2019Angieri \u2013 \u201cThe colored Atlas of Lovebirds\u201d).<\/p>\n<p>&nbsp;<\/p>\n<table>\n<caption>Exemplo 13<\/caption>\n<thead>\n<tr>\n<th><\/th>\n<th colspan=\"2\">p\u00e1lido (lutino parcial)<\/th>\n<th>\u00d7<\/th>\n<th colspan=\"2\">lutino americano<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td><\/td>\n<td colspan=\"2\">Z<sup>ia<\/sup>Z<sup>ia<\/sup><\/td>\n<td>\u00d7<\/td>\n<td colspan=\"2\">Z<sup>i<\/sup>W<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">G:<\/th>\n<td>Z<sup>ia<\/sup><\/td>\n<td>Z<sup>ia<\/sup><\/td>\n<td><\/td>\n<td>Z<sup>i<\/sup><\/td>\n<td>W<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">F1:<\/th>\n<td colspan=\"2\">Z<sup>ia<\/sup>Z<sup>i<\/sup><\/td>\n<td><\/td>\n<td colspan=\"2\">Z<sup>ia<\/sup>W<\/td>\n<\/tr>\n<\/tbody>\n<tfoot>\n<tr>\n<td colspan=\"6\">100% p\u00e1lido (lutino parcial)<\/td>\n<\/tr>\n<\/tfoot>\n<\/table>\n<table>\n<caption>Exemplo 14<\/caption>\n<thead>\n<tr>\n<th><\/th>\n<th colspan=\"2\">p\u00e1lido (lutino parcial) \/ amer.<\/th>\n<th>\u00d7<\/th>\n<th colspan=\"2\">verde<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td><\/td>\n<td colspan=\"2\">Z<sup>ia<\/sup>Z<sup>i<\/sup><\/td>\n<td><\/td>\n<td colspan=\"2\">Z<sup>I<\/sup>W<\/td>\n<\/tr>\n<tr>\n<th scope=\"row\">G:<\/th>\n<td>Z<sup>ia<\/sup><\/td>\n<td>Z<sup>i<\/sup><\/td>\n<td><\/td>\n<td>Z<sup>I<\/sup><\/td>\n<td>W<\/td>\n<\/tr>\n<tr>\n<td><\/td>\n<td>Z<sup>I<\/sup>Z<sup>ia<\/sup><\/td>\n<td>Z<sup>I<\/sup>Z<sup>i<\/sup><\/td>\n<td><\/td>\n<td>Z<sup>ia<\/sup>W<\/td>\n<td>Z<sup>i<\/sup>W<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<table>\n<thead>\n<tr>\n<th><\/th>\n<th>gen\u00f3tipo<\/th>\n<th>fen\u00f3tipo<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<th rowspan=\"2\" scope=\"row\">machos:<\/th>\n<td>Z<sup>I<\/sup>Z<sup>ia<\/sup><\/td>\n<td>verde \/ p\u00e1lido (lutino parcial)<\/td>\n<\/tr>\n<tr>\n<td>Z<sup>I<\/sup>Z<sup>i<\/sup><\/td>\n<td>verde \/ americano<\/td>\n<\/tr>\n<tr>\n<th rowspan=\"2\" scope=\"row\">f\u00eameas:<\/th>\n<td>Z<sup>ia<\/sup>W<\/td>\n<td>p\u00e1lido (lutino parcial)<\/td>\n<\/tr>\n<tr>\n<td>Z<sup>i<\/sup>W<\/td>\n<td>lutino americano<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<h2>Epistasia<\/h2>\n<p>Epistasia \u00e9 o fen\u00f4meno gen\u00e9tico que ocorre quando h\u00e1 o predom\u00ednio (domin\u00e2ncia) de um car\u00e1ter sobre outro cujos genes N\u00c3O s\u00e3o alelos.<\/p>\n<p>Um exemplo nos Agapornis \u00e9 o do fator ino ( alelo americano ) e o fator violeta, situados n\u00e3o s\u00f3 em \u201cloci\u201d mas em cromossomos diferentes. Quando ambos se encontram em homozigose em um mesmo indiv\u00edduo o fen\u00f3tipo lutino prevalece, n\u00e3o se distinguindo o fator violeta, embora a ave em quest\u00e3o seja de direito e de fato um lutino violeta.<\/p>\n<p>Dizemos ent\u00e3o que o fator ino \u00e9 epist\u00e1tico em rela\u00e7\u00e3o ao fator violeta e este por sua vez \u00e9 hipost\u00e1tico \u00e0quele.<\/p>\n<p>&nbsp;<\/p>\n<table>\n<caption>Exemplo<\/caption>\n<thead><\/thead>\n<tbody>\n<tr>\n<td>Z<sup>i<\/sup>Z<sup>i<\/sup>VV<\/td>\n<td rowspan=\"2\">lutinos violetas<\/td>\n<\/tr>\n<tr>\n<td>Z<sup>i<\/sup>WVV<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>Importante observar que violetas lutinos n\u00e3o s\u00e3o visuais. Somente o fator P\u00e1lido sofre intera\u00e7\u00e3o com o fator violeta originando um periquito com urop\u00edgio violeta.<\/p>\n<blockquote><p>&#8220;Epistasia \u00e9 o f\u00eanomeno no qual um car\u00e1ter que seria esperadamente vis\u00edvel se torna n\u00e3o visual podendo este ser dominante ou recessivo em rela\u00e7\u00e3o a outros genes.\u201d<\/p><\/blockquote>\n<h3>Genes \u201cFracos\u201d<\/h3>\n<p>Sabemos que existem alguns fatores que parecem ser delet\u00e9rios. Na verdade parecem proporcionar algum tipo de imunodefici\u00eancia que aumenta a letalidade ou morbidade da muta\u00e7\u00e3o. Assim s\u00e3o os roseicollis fulvos ocidentais (fulvo fosco) e personatus fulvos. H\u00e1 tamb\u00e9m os roseicollis dilu\u00eddos japoneses (golden japon\u00eas) cujas f\u00eameas na cor n\u00e3o conseguem se reproduzir. Na verdade eles s\u00e3o t\u00e3o pouco numerosos que merecem maiores estudos.<\/p>\n<p>Genes letais n\u00e3o s\u00e3o incomuns, por\u00e9m comprovadamente ainda n\u00e3o foram identificados nos Agapornis.<\/p>\n","protected":false},"excerpt":{"rendered":"<p>As Leis de Mendel ou Mendelismo na Cria\u00e7\u00e3o de Agapornis Os princ\u00edpios de gen\u00e9tica que hoje conhecemos e aplicamos em nossa cria\u00e7\u00e3o de Agapornis foram descobertos por um monge de Ordem de Santo Agostinho, nascido em 1822 na Sil\u00e9sia: Gregor Johann Mendel. Mendel, com seu trabalho \u201cExperi\u00eancias com hibridiza\u00e7\u00e3o de plantas\u201d formulou os princ\u00edpios b\u00e1sicos [&hellip;]<\/p>\n","protected":false},"author":1,"featured_media":0,"parent":300,"menu_order":20,"comment_status":"closed","ping_status":"closed","template":"","meta":{"_acf_changed":false,"footnotes":"","_links_to":"","_links_to_target":""},"class_list":["post-320","page","type-page","status-publish","hentry"],"acf":[],"_links":{"self":[{"href":"https:\/\/agasscom.org\/site\/wp-json\/wp\/v2\/pages\/320","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/agasscom.org\/site\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/agasscom.org\/site\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/agasscom.org\/site\/wp-json\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"https:\/\/agasscom.org\/site\/wp-json\/wp\/v2\/comments?post=320"}],"version-history":[{"count":1,"href":"https:\/\/agasscom.org\/site\/wp-json\/wp\/v2\/pages\/320\/revisions"}],"predecessor-version":[{"id":753,"href":"https:\/\/agasscom.org\/site\/wp-json\/wp\/v2\/pages\/320\/revisions\/753"}],"up":[{"embeddable":true,"href":"https:\/\/agasscom.org\/site\/wp-json\/wp\/v2\/pages\/300"}],"wp:attachment":[{"href":"https:\/\/agasscom.org\/site\/wp-json\/wp\/v2\/media?parent=320"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}